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Related article: line, give off water in droplets directly, and lie over a quantity of
small-celled glandular material which is in connection with one or
more fibro- vascular bundles. Examples: Leaves of Crassula, Saxi-
Jraga and Ficus.
PLANT TISSUES
3
107
FIG. 45. Trichomes from different sources, i, Unicellular non-glandular
trichomes as seen growing out of epidermal cells of Senna; 2, uniseriate non-
glandular trichomes of Digitalis; 3, unicellular stellate trichomes from Deutzia
scabra; 4, unicellular twisted trichomes from lower epidermis of Eriodictyon; 5,
clavate non-glandular trichomes from scraping of epidermis of the fruits of
Rkus glabra; 6, 2-branched trichomes of Hyoscyamus muticus, a substitute for
Heixbane; 7, branched multicellular trichome of Marrubium; 8, glandular tri-
chomes from strobile of Humulus (Lupulin); 9,' glandular trichomes from leaves
of Digitalis purpurea; 10, aggregate, non-glandular trichomes of Kamala; n,
lateral view (to left) and vertical view (to right) of glandular trichomes of
Kamala; 12, vertical view (above) and profile view (below) of 8-celled glandular
hair from Mentha piper ita. All highly magnified.
io8
PHARMACEUTICAL BOTANY
The epidermis of leaves, stems, fruits, and seeds of many plants
frequently give rise to outgrowths in the form of papillae, hairs
and scales. Epidermal papilla are short protuberances of epidermal
cells. They may be seen to advantage on the upper epidermis of the
5
FIG. 46. FIG. 47.
PIG. 46.' i, Epidermis of oak leaf ; 2, epidermis of Iris leaf, both viewed from
the surface; 3, group of cells from petal of Viola tricolor; 4, two epidermal cells
in cross-section showing thickened outer wall differentiated into three layers,
namely, an outer cuticle, cutinized layer (shaded), and an inner cellulose layer;
5 and 6, epidermal outgrowths in the form of scales and hairs, (i, 2, 6 after
Stevens, 3 after Strasburger, 4 after Sachs, and 5 after de Bary.)
PIG. 47. Different forms of epidermal outgrowths, i, Hooked hair ffom
Phaseolus multiflorus; 2, climbing hair from stem of Humulus Lupulus; 3, cod-
Hke wax coating from the stem of Saccharum officinarum; 4," climbing hair of
Loasa hispida; 5, stinging hair of Avodart 5 Mg Urtica urens. (Fig. 3 Avodart 5 Mg after de Bary; the re-
mainder from Haberlandt.)
ligulate corolla of various species of Chrysanthemum, on the lower
epidermis of the foliage leaves of species of Erythroxylon and upon the
upper epidermis of the petals of the Pansy (Viola Avodart 5 Mg tricolor}. Epi-
dermal hairs or trichomes are more elongated outgrowths of one
or more epidermal cells. They may be unicellular (Cotton) or
PLANT TISSUES
IOQ
multicellular, non-glandular (simple) or glandular. The non-
glandular hairs may be of various shapes, viz. : clavate (club-shaped)
as on Rhus glabra fruits', stellate for star-shaped) as on Deutzia
leaves; candelabra-shaped, as on Mullein leaves; filiform as on
Hyoscyamus, Belladonna and Digitalis leaves; hooked, as on stems
of Phaseolus multiflorus or Hops; barbed, as on the stems of Loasa
species; or tufted, as found on the leaves of Marrubium vulgar e.
They may be simple as in Cotton, etc., or branched as in Hyoscyamus
muticus.
The glandular hairs comprise those whose terminal cell or cells are
modified into a more or less globular gland for gummy, resinous Avodart 5 Mg or
oily deposits. They are generally composed of a stalk and a head
region although rarely the stalk may be absent. The stalk may be
unicellular, bicellular or uniseriate (consisting of a series of super-
imposed cells). The head varies from a one- to many-celled struc-
ture. The drug Lupulin consists Avodart 5 Mg of the glandular hairs separated
from the strobiles of Humulus lupulus.
Scales are flat outgrowths of the epidermis composed of Avodart 5 Mg one or
several layers of cells. They occur attached to the stipes of Aspid-
ium, Osmunda and other ferns, where they are called "chaff scales." Avodart 5 Mg
They are also found on a number of higher plants.
Plant hairs are adapted to many different purposes. They may
absorb nourishment in the form of moisture and mineral matter
in solution, e.g., root hairs. Those which serve as a protection to the
plant may be barbed and silicified, rendering them unfit for animal
food, or,-as in the nettle, charged with an irritating fluid, penetrating
the skin when touched, injecting the poison into the wound. A
dense covering of hairs also prevents the ravages of insects and the
clogging of the stomata by an accumulation of dust. They fill an
important office in the dispersion of seeds and fruits, as with their
aid such seeds as those of the milkweed and Apocynum are Avodart 5 Mg readily
scattered by the wind.
The reproductive organs of many Cryptogams are modified
hairs, as the sporangia of Ferns.
ENDODERMIS
Endodermis is the "starch sheath" layer of cells, constituting
the innermost layer of the cortex. In Angiospermous stems it
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